Darwinged Fungus Gnat

Introduction

The common name of gnat is applied to certain small flies, and that of fungus which comes from their common occurrence on fungi which serve as a major food source for their larvae; darkwinged describes the smoke-colored wings commonly found within the Sciaridae. These flies are nuisance pests in and around structures, but a few species are agricultural pests. For the Sciaridae, about 137 species, and for the Mycetophilidae, about 714 are found in the United States and Canada.

Recognition

Adults are about 1/23-7/16 inch long; slender to moderately robust, longlegged, somewhat mosquito-like. Color usually black, brown, or yellowish, sometimes brightly colored wings, usually smoke-colored or sometimes patterned with darker areas. Head with eyes separated or touching present. Antenna usually with 16 segments. Wings with front margin thickened to near wing tip; basically and forking beyond middle of wing and cubitus forking in basal quarter of wing; Mycetophilidae with radial sector simple/unbranched or two-branched. Legs with coxae elongated, tibiae with one or two special spurs, pupil absent or minute and narrow to broad but much shorter than claws.

Mature larvae usually are slightly longer than respective adults; sciarid larva with shiny black head and a 12-segmented essentially featureless white translucent body, terminal abdominal segment with a ventral/bottom lobe which acts as a proleg; mycetophillid larva usually cylindrical and slender, with a well-developed head, and 11 or 12 segments, most segments often with entral creeping welts.

Habits

Adults typically are found on or near larval food materials. Larvae fed primarily on fungi growing in the soil and moist decaying organic matter. Sciarid larvae mostly feed on decaying plant material, animal excrement, or fungus, but some feed in rotting wood or under bark of fallen trees. Several sciarid species are economic pests in greenhouses and commercial mushroom houses, and the larvae of the potato.